Seeds are used to produce most commercial crop plants.
However, a significant number of species are propagated by using plant parts
other than seed. By using vegetative parts, the cultivar produced consists of
plants with identical genotypes and is homogeneous. However, the cultivar is genetically
highly heterozygous. Some plant species are sexually reproducing but are propagated
clonally by choice. Such species are improved through hybridization, so that when
hybrid vigor exists it can be fixed and then the improved cultivar propagated
asexually. In seed propagated hybrids, hybrid vigor is highest in the F1, but
is reduced by 50% in each subsequent generation. In other words, whereas
clonally propagated hybrid cultivars may be harvested and used for planting the
next season’s crop without adverse effects, producers of sexually reproducing
species using hybrid seed must obtain a new supply of seed, as previously indicated.
Apomictic cultivars
Apomixis is the phenomenon of production of seed without
the benefit of the union of sperm and egg cells.The seed harvested are thus
genetically identical to the mother plant. Hence, apomictic cultivars have the
same benefits of clonally propagated ones, as previously discussed. In addition,
they have the convenience of vegetative propagation through seed. Apomixis is
common in
perennial forage grasses.
Multilines
Multilines are developed for self-pollinating
species.These cultivars consist of a mixture of specially developed genotypes
called isolines because they differ only
in a single gene. Isolines are developed primarily for disease control, even
though these cultivars,potentially, could be developed to address other
environmental stresses. Isolines are developed by using the techniques of
backcrossing in which the F1 is repeatedly crossed to one of the parents that lacked the gene of interest.
Genetic structure of cultivars and its implications
The products of plant breeding that are released to
farmers for use in production vary in genetic structure and, consequently, the
phenotypic uniformity of the product. Furthermore, the nature of the product
has implications in how it is maintained by the producers regarding the next
season’s planting.
Homozygous and homogeneous cultivars
A cultivar may be genetically homozygous and,
hence, produce a homogeneous phenotype or product. Selfpollinated species are naturally
inbred and tend to be homozygous. Breeding strategies in these species are geared
toward producing cultivars that are homozygous. The products of economic
importance are uniform.
Furthermore, the farmer may save seed from the
current season’s crop for planting the
next season’s crop, without loss of cultivar performance, regarding yield and
product quality. This attribute is especially desirable to producers in many
developing countries where the general tradition is to save seed from the
current season for planting the next season. However, in developed economies
with well-established commercial seed production systems, intellectual property
rights prohibit the re-use of commercial seed for planting the next season’s
crop, thus requiring seasonal purchase of seed by the farmer from seed
companies.
Heterozygous and homogeneous cultivars
The method of breeding of certain crops leaves the cultivar
genetically heterozygous yet phenotypically homogeneous. One such method is
hybrid cultivar production, a method widely used for production of, especially,
outcrossing species such as corn. The heterozygous genetic structure stems from
the fact that a hybrid cultivar is the F1 product of a cross of highly inbred parents.
Crossing such pure lines produces highly heterozygous F1 plants. Because the F1
is the final product released as a cultivar, all plants are uniformly
heterozygous, and hence homogeneous in appearance. However, the seed harvested
from the F1 cultivar is F2 seed, consequently producing maximum heterozygosity and
heterogeneity upon planting. The implication for the farmer is that the current
season’s seed cannot be saved for planting the next season’s crop for obvious
reasons. The farmer who grows hybrid
cultivars must purchase fresh seed from the seed
company for planting each season. Whereas this works well in developed economies,
hybrids generally do not fit well into the farming systems of developing countries
where farmers save seed from the current season for planting the next season’s
crop. Nonetheless,the use of hybrid seed is gradually infiltrating crop
production in developing countries.
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